Geotrupidae – Entomology

id	order	family	subfamily	Tribe	subtribe	Genus	subgenus	Species	subspecies	Author / Year	Evidence no.	origin	type_status	Sex	label_data	orig_description	current_status	references	preservation	photos	notes
id	order	family	subfamily	Tribe	subtribe	Genus	subgenus	Species	subspecies	Author / Year	Evidence no.	origin	type_status	Sex	label_data	orig_description	current_status	references	preservation	photos	notes
1	Coleoptera	Geotrupidae	Bolboceratinae	Bolboceratini		Bolbaffroides		dhofarensis		Král, Hillert, Mencl & Sommer, 2022: 427	P5/COL/99999	general coll.	holotype	♂	OMAN, Dhofar Province / Jabal al Qamar / W Al Mughsayl – Wádí / N 16.84497°E 53.68615° / 20 – 31.8.2012 / lgt. P. Kučera [p] // Bolbaffroides / dhofarensis sp. nov. / HOLOTYPUS ♂ / David Král, Oliver Hillert, / Ladislav Mencl & David / Sommer det. 2021 [p, red label]	KRÁL D., HILLERT O., MENCL L. & SOMMER D. 2022: Bolbaffroides dhofarensis (Coleoptera: Bolboceratidae), a new species from Oman. Zootaxa 5128: 425–434.	valid species		dry; pinned	no
2	Coleoptera	Geotrupidae	Bolboceratinae	Bolboceratini		Bolbaffroides		dhofarensis		Král, Hillert, Mencl & Sommer, 2022: 427	P5/COL/99999	general coll.	allotype	♀	OMAN, Dhofar Province / Jabal al Qamar / W Al Mughsayl – Wádí / N 16.84497°E 53.68615° / 20 – 31.8.2012 / lgt. P. Kučera [p] // Bolbaffroides / dhofarensis sp. nov. / ALLOTYPUS ♀ / David Král, Oliver Hillert, / Ladislav Mencl & David / Sommer det. 2021 [p, red label]	KRÁL D., HILLERT O., MENCL L. & SOMMER D. 2022: Bolbaffroides dhofarensis (Coleoptera: Bolboceratidae), a new species from Oman. Zootaxa 5128: 425–434.	valid species		dry; pinned	no
3	Coleoptera	Geotrupidae	Bolboceratinae	Bolboceratini		Bolbaffroides		dhofarensis		Král, Hillert, Mencl & Sommer, 2022: 427	P5/COL/99999	ex coll. R. Červenka	paratype	♂	OMAN mer. / Darbaat / 18. – 21. 9. 2003 / R. Červenka lgt. [p] // ex coll. R. Červenka / National Museum / Prague, Czech Republic [p] // Bolbaffroides / dhofarensis sp. nov. / PARATYPUS ♂ / David Král, Oliver Hillert, / Ladislav Mencl & David / Sommer det. 2021 [p, red label]	KRÁL D., HILLERT O., MENCL L. & SOMMER D. 2022: Bolbaffroides dhofarensis (Coleoptera: Bolboceratidae), a new species from Oman. Zootaxa 5128: 425–434.	valid species		dry; pinned	yes
4	Coleoptera	Geotrupidae	Bolboceratinae	Bolboceratini		Bolbaffroides		dhofarensis		Král, Hillert, Mencl & Sommer, 2022: 427	P5/COL/99999	general coll.	paratype	♂	S. Oman, W from Salalah / 20km W from Al / Mughsayl h-640m slopes / to Arabian Sea (Camels) / 8-24.7.2007. / Krueger, Saldaitis leg [p] // Bolbaffroides / dhofarensis sp. nov. / PARATYPUS ♂ / David Král, Oliver Hillert, / Ladislav Mencl & David / Sommer det. 2021 [p, red label]	KRÁL D., HILLERT O., MENCL L. & SOMMER D. 2022: Bolbaffroides dhofarensis (Coleoptera: Bolboceratidae), a new species from Oman. Zootaxa 5128: 425–434.	valid species		dry; pinned	no
5	Coleoptera	Geotrupidae	Bolboceratinae	Bolboceratini		Bolbaffroides		dhofarensis		Král, Hillert, Mencl & Sommer, 2022: 427	P5/COL/99999	general coll.	paratype	♂	Sultanate of Oman / Dhofar prov. 21.-31.VIII. / Wadi al Mughsayl / 50 m, 29.-31.VIII.2007 / St. Jakl leg. [p] // Bolbaffroides / dhofarensis sp. nov. / PARATYPUS ♂ / David Král, Oliver Hillert, / Ladislav Mencl & David / Sommer det. 2021 [p, red label]	KRÁL D., HILLERT O., MENCL L. & SOMMER D. 2022: Bolbaffroides dhofarensis (Coleoptera: Bolboceratidae), a new species from Oman. Zootaxa 5128: 425–434.	valid species		dry; pinned	no
6	Coleoptera	Geotrupidae	Bolboceratinae	Bolbochromini		Bolbocerodema		zonatum		Nikolajev, 1973: 859	P5/COL/99999	ex coll. D. Král	paratype	unsexed	Yuzh. Primorie / Kedr. Pad‘ [p] 16.VII. [hw] / L. Zimina [p] 1962 [hw, in Cyrillic script] // Paratypus [p] / Bolbocerodema / zonatum / Nikolajev [hw, red label]	NIKOLAJEV G. V. 1973: Novyy rod i dva novykh vida plastinchatousykh zhukov (Coleoptera, Scarabaeidae, Geotrupinae) iz Palearktiki. (A new genus and two new species of scarabaeid-beetles (Coleopterea, Scarabaeidae, Geotrupinae) from Palaearctics). Entomologicheskoe Obozrenie 52: 856–861 (in Russian, English title).	valid species	BEZBORODOV V. G. & KOSHKIN E. S. 2014: A review of Bolboceratidae (Coleoptera, Scarabaeoidea) species from the Russian Far East. Entomological Review 94: 1313–1319.	dry; pinned; specimen is badly damaged with the head and pronotum missing	no
7	Coleoptera	Geotrupidae	Bolboceratinae	Odonteini		Bolboceras		excavatum		R.A. Philippi in F. Philippi, 1859: 660	P5/COL/99999	ex coll. Nickerl	lectotype	♂	Chile [hw] // Typus [p, red label, black margin] // Coll. Nickerl / Mus. Pragense [p] // Bolboceras / excavatum / Phil. Typ! Chili [hw, green label, black margin] // BOLBOCERAS / EXCAVATUM / PHILIPPI [hw] / LECTOTYPE / A.B.T. SMITH [p, red label] // Southern Neotropical Scarabs / database # AS 2616199 / Bolborhinum laesicolle / (Fairmaire, 1856) ♂ / DET: A.B.T. SMITH 2008 [p]	PHILIPPI F. 1859: Algunas especies nuevas de coleópteros de la provincia de Valdivia. Anales de la Universidad de Chile 16: 656–678.	junior subjective synonym of Bolborrhinum laesicolle (Fairmaire, 1856)	MONCADA J. & SMITH A. B. T. 2008: A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa 1794: 1–48.	dry; pinned	no	SMITH A.B.T. 2015: Rediscovery of primary type specimens and lectotype signation for Bolboceras excavatum R. A. Philippi, 1859 (Coleoptera: Geotrupidae). Acta Entomologica Musei Nationalis Pragae 55: 681–683.
8	Coleoptera	Geotrupidae	Bolboceratinae	Odonteini		Bolboceras		excavatum		R.A. Philippi in F. Philippi, 1859: 660	P5/COL/99999	ex coll. Nickerl	paralectotype	♂	Chile [hw] // Typus [p, red label, black margin] // Coll. Nickerl, Mus. Pragense [p] // BOLBOCERAS / EXCAVATUM / PHILIPPI [hw] / PARALECTOTYPE / A.B.T. SMITH [p, yellow label] // Southern Neotropical Scarabs / database # AS 2616200 / Bolborhinum laesicolle / (Fairmaire, 1856) ♂ / DET: A.B.T. SMITH 2008 [p]	PHILIPPI F. 1859: Algunas especies nuevas de coleópteros de la provincia de Valdivia. Anales de la Universidad de Chile 16: 656–678.	junior subjective synonym of Bolborrhinum laesicolle (Fairmaire, 1856)	MONCADA J. & SMITH A. B. T. 2008: A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa 1794: 1–48.	dry; pinned	no	SMITH A.B.T. 2015: Rediscovery of primary type specimens and lectotype signation for Bolboceras excavatum R. A. Philippi, 1859 (Coleoptera: Geotrupidae). Acta Entomologica Musei Nationalis Pragae 55: 681–683.
9	Coleoptera	Geotrupidae	Bolboceratinae	Odonteini		Bolboceras		excavatum		R.A. Philippi in F. Philippi, 1859: 660	P5/COL/99999	ex coll. Nickerl	paralectotype	♂	Chile [hw] // Typus [p, red label, black margin] // Coll. Nickerl, Mus. Pragense [p]	PHILIPPI F. 1859: Algunas especies nuevas de coleópteros de la provincia de Valdivia. Anales de la Universidad de Chile 16: 656–678.	junior subjective synonym of Bolborrhinum laesicolle (Fairmaire, 1856)	MONCADA J. & SMITH A. B. T. 2008: A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa 1794: 1–48.	dry; pinned	no	SMITH A.B.T. 2015: Rediscovery of primary type specimens and lectotype signation for Bolboceras excavatum R. A. Philippi, 1859 (Coleoptera: Geotrupidae). Acta Entomologica Musei Nationalis Pragae 55: 681–683.
10	Coleoptera	Geotrupidae	Bolboceratinae	Stenaspidiini		Stenaspidius		houstoni		Howden, 1993: 384	P5/COL/99999	general coll.	paratype	unsexed	9-6-’92 / M. Golding [hw] // 28 k. N. Kumarina / W.A. redsand [hw] // PARATYPE / Stenaspidius / houstoni / H. Howden [p, yellow label]	HOWDEN H. F. 1993: New Bolboceratini from Western Australia (Coleoptera: Scarabaeidae: Geotrupinae). Journal of the Australian Entomological Society 32: 379–386.	valid species		dry; pinned	no